Helicase 3D structures

3D structures of helicase3D structures of helicase

Updated on 16-August-2020

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• Helicase Mot1
  • 3oc3 – Hel MOT1 heat domain + TFIID-1 – Encephalitozoon cuniculi
    
• Helicase 308
  • 2p6r – AfHel 308 + DNA – Archaeoglobus fulgidus
    
  • 2p6u - AfHel 308
• DNA helicase
  • 3fld – EcHel I C terminal – Escherichia coli
    
  • 4l0j – EcHel I translocation domain
    
  • 2q7t, 2q7u – EcHel I relaxase domain (mutant)
    
  • 1p4d - EcHel I relaxase domain
    
  • 4n8o, 5n8o - EcHel I + DNA
    
  • 2a0i - EcHel I (mutant) + DNA
    
  • 1e0j, 1e0k – Hel 4D domain – Phage T7
• DNA helicase II or UvrD
  • 1q2z, 1rw2 – hHel II KU86 C terminal – human - NMR
    
  • 5y58 – yHel II + DNA – yeast
    
  • 5y59 – yHel II subunit 2 + Sir4P peptide
    
  • 3lfu – EcHel II
    
  • 2is1, 2is2 - EcHel II (mutant) + DNA
    
  • 2is4, 2is6 - EcHel II (mutant) + ADP + DNA
    
  • 4c2t, 4c2u, 4c30 – DrHel II + DNA - Deinococcus radiodurans
    
  • 6ppj – MsHel (mutant) – Mycobacterium smegmatis – Cryo EM
    
  • 6ppr, 6ppu – MsHel + DNA – Cryo EM
    
• ATP-dependent helicase ATRX
  • 3qln - hATRX ADD domain residues 167-289 – human
    
  • 2ld1, 2jm1 - hATRX ADD domain - NMR
    
  • 3ql9, 4w5a – hATRX ADD domain + histone H3 peptide
    
  • 2lbm - hATRX ADD domain + histone H3 peptide – NMR
    
  • 3qla, 3qlc - hATRX ADD domain (mutant) + histone H3 peptide
    
  • 5y6o – hATRX N terminal + ATRX
    
  • 2xzp - hRENT1 helicase domain
    
  • 2wjv - hRENT1 helicase CH and C terminal domains
    
  • 2wjy - hRENT1 helicase CH domains
    
  • 2xzo – hRENT1 helicase domain + RNA
    
• ATP-dependent helicase Fun30
  • 5gn1 – yFun C-terminal
    
• ATP-dependent helicase Rho
  • 3l0o – TmRho – Thermotoga maritima
    
  • 3ice – EcRho + ADP + BeF3 + RNA
    
  • 2ht1 - EcRho + RNA
    
  • 1a8v, 1a62 – EcRho RNA-binding domain
    
  • 1a63 - EcRho RNA-binding domain – NMR
    
  • 2a8v – EcRho RNA-binding domain + RNA
    
  • 1pv4 – EcRho + DNA
    
  • 1pvo – EcRho + ssRNA + ANPPNP
    
  • 1xpr, 1xpu, 1xpo - EcRho + ssRNA + antibiotic
• ATP-dependent helicase MFD
  • 3hjh – EcMFD motor domain
• ATP-dependent helicase HEPA
  • 6bog – EcHEPA
• ATP-dependent helicase E1
  • 2v9p – PvE1 helicase domain – Bovine papillomavirus
    
  • 2gxa – PvE1 + DNA + Mg-ADP
    
  • 1tue – PvE1 + E2 – human papillomavirus
• ATP-dependent helicase CHD
  • 2b2y – hCHD1
    
  • 4b4c – hCHD1 DNA-binding domain
    
  • 2b2t, 2b2u, 2b2v, 2b2w – hCHD1 (mutant) + histone H3 tail
    
  • 4nw2 – hCHD1 + NS1 peptide
    
  • 2ee1 – hCHD4 chromodomain – NMR
    
  • 2l5u – hCHD4 PHD1 domain – NMR
    
  • 2l75 – hCHD4 PHD2 domain + histone H3 peptide– NMR
    
  • 2epb – hCHD6 chromodomain – NMR
    
  • 2ckc, 2v0e, 2v0f – hCHD7 BRK domain - NMR
    
  • 2dl6 – hCHD8 BRK domain - NMR
    
  • 2cka – hCHD8
    
  • 2h1e, 2xb0, 3mwy – yCHD1 chromodomain
    
  • 2dy7 - yCHD1 chromodomain 1 – NMR
    
  • 2dy8 - yCHD1 chromodomain 2 – NMR
    
  • 3ted - yCHD1 chromodomain 1 + DNA
    
• ATP-dependent DNA helicase PIF1

• • 5fhh, 6hpt – hPIF residues 200-641
    
  • 6hpu, 6hpq, 6hph – hPIF residues 206-620 + nucleotide
    
  • 5o6b, 5o6d – yPIF residues 237-780 + DNA + ATP
    
  • 5o6e – yPIF residues 237-780 + DNA + ADP
    
• ATP-dependent DNA helicase TA0057

• • 5h8w – TaTA0 + DNA – Thermoplasma acidophilum
    
• ATP-dependent DNA helicase LHR

• • 5v9x – MsLHR + DNA – Mycobacterium smegmatis
    
• Dead box ATP-dependent RNA helicase
  • 4xw3 – hDDX1 SPRY domain
    
  • 2i4i – hDDX3X residues 167-581
    
  • 4px9 - hDDX3X residues 135-407
    
  • 4pxa - hDDX3X residues 135-407 (mutant)
    
  • 5e7i - hDDX3X residues 133-584
    
  • 2jgn – hDDX3X helicase domain residues 408-579
    
  • 3ly5 – hDDX18 dead domain
    
  • 6l5l, 6l5o – hDDX21 residues 120-495 (mutant)
    
  • 6l5m – hDDX21 + AMP
    
  • 6l5n – hDDX21 + RNA
    
  • 2rb4 – hDDX25 helicase domain
    
  • 2p6n – hDDX41 helicase domain (mutant)
    
  • 2e29, 2m3d - hDDX50 GUCT domain - NMR
    
  • 1vec – hP54 N terminal
    
  • 3fho – DBP5 – fission yeast
    
  • 2kbe – yDBP5 N terminal – NMR
    
  • 2kbf - yDBP5 C terminal – NMR
    
  • 3gfp - yDBP5 C-terminal
    
  • 2hjv – BsDBPA domain 2 – Bacillus subtilis
    
  • 2g0c - BsDBPA RNA-binding domain
    
  • 3eaq, 3ear, 3eas – TtHera residues 215-426 - Thermus thermophilus
    
  • 3i31 - TtHera residues 431-517
    
  • 3i32 - TtHera residues 218-517
    
  • 3nej - TtHera N terminal (mutant)
    
  • 5ivl – GsDB CSHA - Geobacillus stearothermophilus
    
  • 5b88 - EcDBP RRM-like domain - NMR
    
  • 5gju - EcDBP residues 6-210
    
  • 5gi4 - EcDBP residues 218-445
    
  • 6aib – SaDB CSHA N-terminal – Staphylococcus aureus
    
• Dead box ATP-dependent RNA helicase complexes
  • 3fmo, 3fhc – hDBP5 + Nup214
    
  • 3fht – hDBP5 + AMPPNP + RNA
    
  • 6cz5 - hDDX3X residues 132-607 + AMP
    
  • 6o5f - hDDX3X + RNA
    
  • 6b4k – hDDX19B residues 54-479 + ANP
    
  • 6b4j, 6b4i – hDDX19B residues 54-479 + GLE1 + Nup42 + ANP
    
  • 3fmp – hDDX19B + Nup214
    
  • 3g0h – hDDX19B residues 54-275 + ATP analog + RNA
    
  • 3ews - hDDX19B residues 54-275 + ADP
    
  • 5e7m, 5e7j – hDDX3X catalytic domain + nucleotide
    
  • 3dkp – hDDX52 domain I + ADP
    
  • 2wax, 2way – hDDX6 C terminal + EDC3-FDF peptide
    
  • 2j0s, 2j0q, 2hyi – hDDX48 + protein mago nashi homolog + RNA-binding protein 8A + protein CASC3 + RNA
    
  • 2j0u - hDDX48 + protein CASC3
    
  • 2db3 – DmVASA residues 200-623 + RNA – Drosophila melanogaster
    
  • 5nt7 – DmVASA RecA-like domain + maternal effect protein
    
  • 3rrm – yDBP5 residues 91-482 (mutant) + GLE1 + NUP159 + ADP + IP6
    
  • 3rrn - yDBP5 residues 91-482 (mutant) + GLE1 + IP6
    
  • 5elx - yDBP5 residues 91-482 + RNA
    
  • 3pew, 3pey - yDBP5 residues 91-482 (mutant) + ADP + BeF3 + RNA
    
  • 3peu, 3pev - yDBP5 C-terminal (mutant) + GLE1 + IP6
    
  • 3i5x, 3i5y – yMSS116 + AMPPNP + RNA
    
  • 3sqw, 3sqx - yMSS116 + AMPPNP + RNA
    
  • 3i61, 4tyw, 4tyy, 4tz0, 4tz6 - yMSS116 + nucleotide + BeF3 + RNA
    
  • 3i62 - yMSS116 + ADP + AlF4 + RNA
    
  • 4tyn - yMSS116 + ADP + DNA
    
  • 2xau, 3kx2, 5jpt – yPrp43 + nucleotide
    
  • 5i8q - yPrp43 + ADPNP + RNA
    
  • 5ltk, 5ltj, 5d0u - CtPrp43 + ADP – Chaetomium thermophilum
    
  • 5lta - CtPrp43 + ADP + RNA
    
  • 3jrv – DDX3X + protein K7 – Vaccinia virus
    
  • 3nbf - TtHera N terminal (mutant) + 8-oxo-ADP
    
  • 3moj - BsDBPA RNA-binding domain + RNA
    
  • 6aic – SaDB CSHA N-terminal + AMP
    
• DEAH box RNA helicase
  • 2eqs – hDHX8 S1 domain residues 260-355
    
  • 3i4u – hDHX8 residues 950-1183
    
  • 6hys – hDHX8 helicase domain residues 548-1220 + ATP
    
  • 6hyt – hDHX8 helicase domain + ADP
    
  • 6hyu – hDHX8 helicase domain + RNA
    
  • 3llm – hDHX9
    
  • 5xdr – hDHX15 residues 110-795 (mutant)
    
  • 6sh6, 6sh7 – hDHX15 + Nfkb-repressing factor + ADP
    
  • 2n16 – hDHX36 G-quadruplex domain - NMR
    
  • 2rqa – hDHX58 C terminal – NMR
    
  • 3vyx, 3vyy – hDHX9 RNA-binding domain + RNA
    
  • 2n21 – hDHX36 G-quadruplex domain + DNA - NMR
    
  • 5e3h - hDHX58 residues 232-925 + RNA
    
  • 3eqt- hDHX58 residues 541-678 + RNA
    
  • 5vha – bDHX36 residues 150-1010 - bovine
    
  • 5vhc, 5vhd – bDHX36 residues 150-1010 + ADP
    
  • 5vhe – bDHX36 residues 56-1010 + DNA
    
  • 2rs6, 2rs7 – mDHX9 RNA-binding domain – NMR
    
  • 6heg, 6eud – EcHRPB
    
• ATP-dependent RNA helicase
  • 2g9n - hEIF4A dead domain
    
  • 5zbz - hEIF4A dead domain + sanguinarine
    
  • 5zc9 - hEIF4A dead domain + ATP + RNA
    
  • 6ieg - hDOB1 residues 70-1042
    
  • 6ro1 - hDOB1 residues 70-1042 + NVL
    
  • 6ieh - hDOB1 residues 70-1042 + NRDE2
    
  • 6c90 - hDOB1 residues 70-600 + ZCCHC8
    
  • 6d6r, 6d6q - hDOB1 in exosome + RNA – Cryo EM
    
  • 2xzl – yNAM7 CH and helicase domains + RNA
    
  • 2xgj – yDOB1 residues 81-1073 + RNA
    
  • 4u4c - yDOB1 residues 81-1073 + poly(A) RNA polymerase protein 2
    
  • 4qu4 - yDOB1
    
  • 5ooq - yDOB1 residues 81-1073 + NOP53
    
  • 5suq – ySUB2 + TEX1 + THO2
    
  • 5sup – ySUB2 + YRA1 + RNA
    
  • 2vso, 2vsx – yEIF4A + initiation factor 4F middle domain
    
  • 2g9n - hEIF4A dead domain
• Viral ATP-dependent RNA helicase NS3
  • 3o8b, 3o8d, 1cu1, 8ohm, 1hei, 4a92, 3rvb – HvHel NS3 residues 1186-1658 – Hepatitis C virus
    
  • 1onb, 1jr6 - HvHel NS3 arginine-rich domain residues 1353-1507 – NMR
    
  • 2jlq, 2bhr, 2bmf, 2whx, 2wzq – DvHel NS3 residues 1646-2092 – Dengue virus
    
  • 5xc7 – DvHel NS3 residues 1646-2092 (mutant)
    
  • 2z83 – Hel NS3 residues 167-624 – Japanese encephalitis virus
    
  • 2v8o, 2wv9 - Hel NS3 helicase domain – Murray valley encephalitis virus
    
  • 2qeq - Hel NS3 residues 1691-2124 – Kunjin virus
    
  • 5txg, 5jmt, 5vi7, 5jps, 6mh3 - ZvHel NS3 helicase domain – Zika virus
    
  • 5k8u, 5k8l, 5k8i, 5jwh, 5jrz - ZvHel NS3 residues 164-621
    
  • 5x8y - ZvHel NS1 residues 168-352
    
  • 2v6i – KvHel helicase domain – Kokobera virus
    
  • 2v6j - KvHel helicase domain (mutant)
    
  • 5wso, 5gvu - Hel NS3 residues 1939-2414 – bovine viral diarrhea virus
    
• Viral ATP-dependent RNA helicase NS3 complexes
  • 3o8c, 3o8r, 3kqh, 3kqk, 3kql, 3kqn, 3kqu, 1a1v - HvHel NS3 + RNA
    
  • 2f9v - HvHel NS3 protease domain + polyprotein
    
  • 2a4g, 1rtl, 1w3c, 1dxp - HvHel NS3 protease domain + NS4A peptide
    
  • 2a4q, 2a4r, 2f9u, 2fm2, 1dy8, 1dy9 - HvHel NS3 protease domain + NS4A peptide + inhibitor
    
  • 2jlr - DvHel NS3 residues 1646-2092 + AMPPNP
    
  • 2jls - DvHel NS3 residues 1646-2092 + ADP
    
  • 2jlu, 2jlw, 5xc6 - DvHel NS3 residues 1646-2092 + RNA
    
  • 2jlv - DvHel NS3 residues 1646-2092 + AMPPNP + RNA
    
  • 2jlx, 2jly, 2jlz - DvHel NS3 residues 1646-2092 + ADP + RNA
    
  • 2vbc - DvHel NS3 residues 1475-2092 + NS2A peptide
    
  • 5k8t - ZvHel NS3 residues 164-621 + GTP
    
  • 5gjc - ZvHel NS3 residues 1676-2119 + ATP
    
  • 5gjb - ZvHel NS3 residues 1676-2119 + RNA
    
  • 5y4z - ZvHel NS3 residues 1676-2119 + AMPPNP
    
• ATP-dependent RNA helicase SUV3
  • 3rc3 – hSUV3 residues 47-722
    
  • 3og8 – hDDX58 C terminal + RNA
    
  • 3rc8 - hSUV3 residues 47-722 + RNA
    
  • 6f4a – ySUV3 + exoribonuclease II + RNA
    
• ATP-dependent RNA helicase A
  • 3llm – hHel A
    
  • 3vyx, 3vyy - hHel A RNA-binding domain + RNA
    
  • 1whq, 2rs6, 2rs7 – Hel A RNA-binding domain – mouse - NMR
    
• ATP-dependent RNA helicase Repa
  • 1uaa – EcRepa + DNA
    
  • 1nlf - EcRepa + sulfate
    
  • 1g8y, 1olo - EcRepa
• ATP-dependent RNA helicase HEF
  • 1x2i – HEF DNA-binding domain – Pyrococcus furiosus
• ATP-dependent RNA helicase Srmb
  • 2yjt - EcSrmb residues 219-388 + RRAA
    
• ATP-dependent RNA helicase Dhh1
  • 4bru – yDhh1 (mutant) + EDC3
    
  • 4brw – yDhh1 (mutant) + PAT1
    
• Werner syndrome ATP-dependent DNA helicase
  • 2e1e, 2e1f, 2dgz – hWRN HRDC domain 1142-1242
    
  • 2fbt - hWRN exonuclease domain 38-236
    
  • 2fbv, 2fbx, 2fby, 2fc0 - hWRN exonuclease domain + metal ion
  • 3aaf – hWRN C terminal 949-1079 + DNA
    
  • 6yhr - hWRN 517-1094
    
  • 2e6l, 2e6m - mWRN exonuclease domain
• ATP-dependent DNA helicase Q
  • 2v1x - hQ1 residues 49-616
    
  • 2wwy, 4u7d – hQ1 residues 49-616 + DNA
    
  • 5lst – hQ4 residues 427-1116
    
  • 2kmu – hQ4 N terminal - NMR
    
  • 5lb8 – hQ5
    
  • 5lb5, 5lb3 – hQ5 + ADP
    
  • 5lba – hQ5 + urea derivative
    
• ATP-dependent DNA helicase RecG
  • 1gm5 – TmRecG + DNA
    
• ATP-dependent DNA helicase RecQ (Bloom Syndrome helicase)
  • 4cgz, 4o3m - hRecQ catalytic domain + DNA
    
  • 4cdg - hRecQ catalytic domain + nanobody
    
  • 1d8b - yRecQ HRDC domain - NMR
    
  • 3iuo – RecQ residues 604-725 – Porphyromonas gingivalis
    
  • 2rhf – DrRecQ HRDC domain 3 – Deinococcus radiodurans
    
  • 2ma1 - DrRecQ HRDC domain – NMR
    
  • 4q48 - DrRecQ catalytic domain
    
  • 4q47 - DrRecQ catalytic domain + ADP
    
  • 1wud - EcRecQ HRDC domain
    
  • 1oyw - EcRecQ catalytic domain
    
  • 1oyy - EcRecQ catalytic domain + ATP
    
• ATP-dependent DNA helicase RuvA, RuvB
  • 2ztc, 2ztd, 2zte, 2h5x – MtRuvA – Mycobacterium tuberculosis
    
  • 1bvs - RuvA – Mycobacterium leprae
    
  • 1cuk, 1hjp – EcRuvA
    
  • 1d8l – EcRuvA N terminal
    
  • 1c7y, 1bdx – EcRuvA + DNA
    
  • 1hqc – TtRuvB
    
  • 1ixr, 1ixs – TtRuvA + RuvB (mutant)
    
  • 1in4 – TmRuvB
    
  • 1in5, 1in6, 1in7, 1in8, 1j7k – TmRuvB (mutant)
    
  • 3pfi – RuvB + ADP – Campylobacter jejuni
    
  • 6blb - PaRuvB + ADP - Pseudomonas aeruginosa
    
• ATP-dependent DNA helicase RuvB-like
  • 2c9o – hRuvBL1 + ADP
    
  • 6h7x – hRuvBL2
    
  • 3uk6 – hRuvBL2 + ADP
    
  • 2cqa – hRuvBL2 C-terminal - NMR
    
  • 2xsz – hRuvBL1 + hRuvBL2 + ATP
    
  • 6qi8, 6qi9 – hRuvBL1 + hRuvBL2 + ADP
    
  • 5oaf – hRuvBL1 + hRuvBL2 + ADP - Cryo EM
    
  • 6fo1 – hRuvBL1 + hRuvBL2 + RPAP3 + ADP - Cryo EM
    
  • 6igm – hRuvBL1 + hRuvBL2 + actin-related protein + helicase SRCAP + protein - Cryo EM
    
  • 6hts – hRuvBL1 + hRuvBL2 + INO80 + ARP5 + histones + DNA - Cryo EM
    
  • 5oun – yRuvBL2 DII domain - yeast - NMR
    
  • 6gej, 6gen – yRuvBL1 + yRuvBL2 + vacuolar protein sorting-associated proteins + ARP6 + SWR complex protein 6 + histones + helicase SWR1 + ADP + ATP + DNA - Cryo EM
    
  • 4wvy – CtRuvBL1 + CtRuvBL2 + ATP
    
  • 4ww4 – CtRuvBL1 + CtRuvBL2 + ADP
    
  • 6fhs – CtRuvBL1 + CtRuvBL2 + INO80 + LES2 + LES5 + ARP5 + ADP + ATP - Cryo EM
    
  • 6fml – CtRuvBL1 + CtRuvBL2 + INO80 + LES2 + LES6 + actin-related protein + histones + ADP + ATP + DNA - Cryo EM
    
  • 5f4h, 5yww – RuvBL (mutant) - Sulfolobus islandicus
    
• ATP-dependent DNA helicase UVSW
  • 2oca, 1rif- T4UVSW (mutant) – Enterobacteria phage T4
    
  • 2jpn - T4UVSW
• ATP-dependent DNA helicase DDA
  • 3upu – T4DDA + DNA
    
• ATP-dependent DNA helicase PriA
  • 2d7e – EcPriA DNA-binding domain
    
  • 2d7g, 2d7h, 2dwl, 2dwm, 2dwn – EcPriA N terminal + polynucleotide
• ATP-dependent DNA helicase PcrA
  • 1qhg – GsPcrA (mutant) + ADPNP
    
  • 1qhh - GsPcrA + ADPNP
    
  • 2pjr, 3pjr - GsPcrA + DNA
    
  • 1pjr - GsPcrA
    
  • 5dma – GsPcrA C-terminal tudor domain
    
• DnaB helicase or replicative DNA helicase
  • 3gxv – HpDnaB – Helicobacter pylori
    
  • 4a1f – HpDnaB C terminal
    
  • 2r6d – BsDnaB – Bacillus stearothermophilus
    
  • 2r6e, 2r6d - GsDnaB
    
  • 2r5u – MtDnaB N terminal
    
  • 3bgw – BpDnaB-like – Bacillus phage SPP1
    
  • 3bh0 - BpDnaB-like G40P ATPase domain
    
  • 2q6t – DnaB – Thermus aquaticus
    
  • 1mi8, 6frh – SyDnaB – Synechocystis
    
  • 6frg, 6fre – SyDnaB (mutant)
    
  • 6bbm – EcDnaB + replication protein P – Cryo EM
    
  • 1b79 - EcDnaB N terminal
    
  • 1jwe - EcDnaB N terminal - NMR
    
  • 2r6c, 2r6a – BsDnaB + DnaG helicase-binding domain
    
  • 6bs8, 6own – MsDnaB 238-376
    
  • 6rix, 6riy – DnaB (mutant) – Mycobacterium chimaera
    
• DnaC helicase
  • 2vyf– GkDnaC – Geobacillus kaustophilus
    
  • 2vye – GkDnaC + DNA
    
  • 3ec2 – AaDnaC – Aquifex aeolicus
    
  • 3ecc – AaDnaC + ADP
    
  • 6qel – EcDnaC + replicative DNA Hel
    
  • 6qem – EcDnaC + replicative DNA Hel + DNA
    
• Helicase BLM
  • 5lup – hBLM DHBN domain
    
  • 5lut – BLM DHBN domain - chicken
    
  • 5lus – BLM DHBN domain - pelican
    
• Helicase CA8 or Prp28
  • 4w7s – yCa8
    
• Helicase NSP
  • 6jim – NSP2 + RNA – Chicungunya virus
    
  • 2hwk – NSP2 – Venzuelan equine encephalitis virus
    
  • 6jyt – NSP13 – Human SARS-COV virus
    
• Helicase SEN1
  • 5mzn, 6i59 – ySEN1
    
• Helicase DING
  • 6fws, 6fwr – EcDING + DNA + ADP
    
• Helicase SNF2
  • 1z5z – SsSNF2 C terminal – Sulfolobus solfataricus
    
  • 1z63 - SsSNF2 ATPase core + DNA
    
  • 1z6a - SsSNF2 ATPase core
• Helicase SKI2
  • 4a4k – ySKI2 insertion domain
    
  • 4a4z – ySKI2 + AMPPNP
    
  • 2va8 – SsSKI2
    
• Helicase SWR1
  • 5i9e – ySWR1 + actin + actin-related protein
    
• Helicase XPD
  • 3crv, 3crw, 5h8c – XPD – Sulfolobus acidocaldarius
    
  • 4a15 - TaXPD + DNA
    
  • 2vsf - TaXPD
    
  • 2vl7 - XPD - Sulfolobus tokodaii
    
  • 5ivw - hXPD + XPB + TFIIH ubunits 2,3,4,5 + SCP-X + SCP-Ybr />
• Pre-mRNA-splicing Helicase
  • 3hib, 3im1, 3im2 – yBrr2 2nd SEC63 domain
    
  • 4bgd, 5m5p, 5m52, 5dca – yBrr2 + pre-mRNA-splicing factor 8
    
• crispr-associated helicase Cas3
  • 4q2c – TteCas3 – Thermobaculum terrenum
    
  • 4q2d – TteCas3 + DTP
    
  • 4qqw, 4qqx – TfCas3 – Thermobifida fusca
    
  • 4qqy – TfCas3 + ADP + HD nuclease
    
  • 4qqz – TfCas3 + AMPPNP
    
  • 6c66 – TfCas3 + Cse1 + Cse3 + Cse4 + Cas5e – Cryo EM
    
  • 5gqh – PaCas3 + anti-crispr protein 3
    
  • 5b7i – PaCas3 + ACRF3